Lassie is one of Hollywood’s most iconic animal actors, originating from Eric Knight’s 1940 novel Lassie Come-Home and the 1943 MGM film. The role was first played by Pal, a male Collie trained by Rudd Weatherwax, despite the character being female—male Collies were preferred for their striking appearance. Pal and his descendants continued the role in films and the legendary 1954–1973 TV series, which made Lassie a global symbol of loyalty and bravery. Generations of Pal’s offspring have kept the tradition alive, cementing Lassie’s place as one of entertainment’s most beloved stars.